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Initially distributed with the PAL release of Need for Speed Undercover
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IOS55 is an IOS that only two games appear to use -- "Need for Speed: Undercover" and "Sonic and the Black Knight".
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It can be an update for the new wireless USB Driving Wheel from Logitech, supported only by Need for Speed: Undercover.
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This IOS is presumably required by Undercover to make a Logitech USB steering wheel work -- however, this does not explain why Sonic and the Black Knight requires it. If a game that requires this IOS is started without it (via brickblocker or a Starfall update skip, for example), the screen will turn blue and a white-text error message "Error #002" will appear.
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Installation from NfS Undercover-disc keeps [[System Menu 3.3]] and does not(!) install [[System Menu 3.4]] when system had 3.3 before.
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Installation from the Undercover disc keeps [[System Menu 3.3]] and does not(!) install [[System Menu 3.4]], assuming the user's Wii didn't already have 3.4.